Board Thread:General Discussion/@comment-26830922-20170211043007/@comment-30535075-20170212222109

The Hollows have Human souls, and the scan specifies that no matter who dies, their soul will go to Yhwach. By spreading pieces of his soul, he meant the Quincies. Not actual physical souls, we didn't see.

As shown with Masaki and Katagiri you don't need to drink Yhwach's blood, to get killed by Yhwach's soul manipulatiom abilities..